PUBLICATIONS
Williams, C.J., Pani, M., Bucchi, A., Smith, R.E., Kao, A., Keeble, W., Ibrahim, N., Martill, D.M. (2021). Helically arranged cross struts in azhdarchid pterosaur cervical vertebrae: implications for the feeding behaviour of giant pterosaurs. iScience.
Beevor, T., Quigley, A., Smith, R., Smyth, R.S.H., Ibrahim, N., Martill, D.M. (2020). Taphonomic evidence supports an aquatic lifestyle for Spinosaurus. Cretaceous Research.
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Shears Ozeki, C., Martill, D.M., Smith, R., Ibrahim, N. (2020). Biological modification of bones in the Cretaceous of North Africa. Cretaceous Research.
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Ibrahim, N., Maganuco, S., Dal Sasso, C., Fabbri, M., Auditore, M., Bindellini, G., Martill, D.M., Zouhri, S., Mattarelli, D., Unwin, D.M., Wiemann, J., Bonadonna., D., Amane, A., Jakubczak, J., Joger, U., Lauder, G.V., Pierce, S.E. (2020). Tail-propelled aquatic locomotion in a theropod dinosaur. NATURE.
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Smyth, R.S.H., Ibrahim, N., Martill, D.M. (2020). Sigilmassasaurus is Spinosaurus: a reappraisal of African spinosaurids. Cretaceous Research.
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Martill, D.M., Smith, R., Unwin, D., Kao, A., McPhee, J., Ibrahim, N. (2020). A new tapejarid (Pterosauria, Azhdarchoidea) in Africa. Cretaceous Research.
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McPhee, J., Ibrahim, N., Kao, A., Unwin, D.M., Smith, R., Martill, D.M. (2020). A new ?chaoyangopterid (Pterosauria: Pterodactyloidea) from the Cretaceous Kem Kem beds of Southern Morocco. Cretaceous Research.
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Jacobs, M.L., Martill, D.M., Unwin, D.M., Ibrahim, N., Zouhri, S., Longrich, N. (2020) Toothed pterosaurs (Pterosauria: Ornithocheiridae) of the mid Cretaceous Kem Kem beds of Morocco: implications for pterosaur paleobiogeographic distribution in the Early Cretaceous. Cretaceous Research.
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Ibrahim, N., Sereno, P.C., Varricchio, D.J., Martill, D.M., Dutheil, D.B., Unwin, D.M., Baidder, L., Larsson, H.C.E., Zouhri, S., Kaoukaya, A. (2020). Geology and paleontology of the Upper Cretaceous Kem Kem Group of eastern Morocco. ZooKeys.
Smyth, R.S.H., Ibrahim, N., Kao, A., Martill, D.M. (2020). Abelisauroid cervical vertebrae from the Cretaceous Kem Kem beds of Southern Morocco and a review of Kem Kem abelisauroids. Cretaceous Research.
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Jacobs, M.L., Martill, D.M., Ibrahim, N., Longrich, N. (2019) A new species of Coloborhynchus (Pterosauria: Ornithocheiridae) from the mid-Cretaceous of North Africa. Cretaceous Research, 84, 1-12.
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Dahdul, W., Manda, P., Cui, H., Dececchi, T.A., Ibrahim, N., Lapp, H., Vision, T., Mabee, P. (2018) Annotation of phenotypes using ontologies: a Gold Standard for the training and evaluation of natural language processing systems. Database, https://doi.org/10.1093/database/bay110
Martill, D.M., Ibrahim, N., Bouaziz, S. (2018) A giant pterosaur in the Early Cretaceous (Albian) of Tunisia. Journal of African Earth Sciences, 147, 331-337.
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Martill, D.M., Unwin, D.M., Ibrahim, N., Longrich, N. (2017) A new and unusual edentulous pterosaur from the Cretaceous Kem Kem beds of North Africa. Cretaceous Research, 84, 1-12.
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Ibrahim, N., Zouhri, S., Sereno, P.C. (2017) Les dinosaures du Maroc – aperçu historique et travaux récents. In ''Paléontologie des Vertébrés du Maroc '', Samir Zouhri (Ed.). Société géologique de France, Mémoire 180.
Klein, C., Longridge, N., Ibrahim, N., Zouhri, S., Martill, D.M. (2017) A new basal snake from the mid-Cretaceous of Morocco. Cretaceous Research http://dx.doi.org/10.1016/j.cretres.2016.12.001
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Ibrahim, N., Dal Sasso, C., Maganuco, S., Fabbri, M., Martill, D.M., Gorscak, E., Lamanna, M.C., (2016) Evidence of a derived titanosaurian (Dinosauria, Sauropoda) in the ‘Kem Kem beds’ of Morocco, with comments on sauropod paleoecology in the Cretaceous of Africa. In ''Recent advances in Cretaceous biodiversity, paleoenvironments and palaeobiogeographic implications'', Ashu Khosla, Spencer Lucas (Eds.). New Mexico Museum of Natural History and Science Bulletin.
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Cui, H., Dahdul, W., Dececchi, T.A., Ibrahim, N., Mabee, P., Balhoff, J.P., Gopalakrishnan, H. (2015) CharaParser+EQ: Performance Evaluation without Gold Standard. Association for Information Science and Technology Annual Meeting.
Dahdul, W., Dececchi, T.A., Ibrahim, N., Lapp, H., Mabee, P. (2015) Moving the mountain: Analysis of the effort required to transform comparative anatomy into computable anatomy. Database, 1-7, doi: 10.1093/database/bav040
Martill, D.M., Ibrahim, N. (2015). An unusual modification of the jaws in cf. Alanqa, a mid-Cretaceous azhdarchid pterosaur from the Kem Kem beds of Morocco. Cretaceous Research, 53, 59-67.
Deans, A.R., Lewis, S.E., Huala, E., Anzaldo, S. S., [...] Ibrahim, N.,[...] Woolley, J.B., Yoder, M. J., Zorn, A. M., Mabee, P.M.,. (2015) Finding our way through Phenotypes. PLOS Biology, PLoS Biol 13(1): e1002033. doi:10.1371/journal.pbio.1002033
Ibrahim, N., Sereno, P. C., Dal Sasso, C., Maganuco, S., Fabbri, M., Martill, D. M. Zouhri, S., Myhrvold, N., Iurino, D. A. (2014). Semiaquatic adaptations in a giant predatory dinosaur. SCIENCE, 345(6204), 1613-1616.
Ibrahim, N., Varricchio, D.J., Sereno, P.C., Wilson, J.A., Dutheil, D.B., Martill, D.M., Baidder, L., Zouhri, S. (2014) Dinosaur footprints and other ichnofauna from the Cretaceous Kem Kem beds of Morocco. PLoS ONE 9(3): e90751 doi:10.1371/journal.pone.0090751.
Haendel, M.A., Balhoff, J.P., Bastian, F., Blackburn, D., Blake, J.A., Comte, A., Dahdul, W., Dececchi, T.A., Druzinsky, R.E., Hayamizu, T.F., Ibrahim, N., Lewis, S.E., Mabee, P.M., Niknejad, A., Robinson-Rechavi, M., Sereno, P.C., Mungall C.J. (2013) Uberon: Unification of multi-species vertebrate anatomy ontologies for comparative biology. Journal of Biomedical Semantics, 5: 21.
Midford, P.E., Balhoff, J.P., Blackburn, D., Dahdul, W., Dececchi, T.A.,Ibrahim, N., Kothari, C.R., Lapp, H., Lundberg, J.G., Mabee, P.M., Sereno, P.C., Vision, T.J., Westerfield, M. (2013) The Vertebrate Taxonomy Ontology: A framework for reasoning across model organism and species phenotypes. Journal of Biomedical Semantics, 4: 34.
Ibrahim N., Kutschera U. (2013). The ornithologist Alfred Russel Wallace and the controversy surrounding the dinosaurian origin of birds. Theory in Biosciences, 132, 267-275.
Martill, D.M., Ibrahim, N., 2012. Aberrant rostral teeth of the sawfish Onchopristis numidus from the Kem Kem beds (?early Late Cretaceous) of Morocco and a reappraisal of Onchopristis in New Zealand. Journal of African Earth Sciences, 64, 71-76.
Martill D.M., Ibrahim N., Brito P.M., Baidder L., Zouhri S., Loveridge R., Naish D. and Hing R. (2011). A new Plattenkalk Konservat Lagerstätte in the Upper Cretaceous of Gara Sbaa, south-eastern Morocco. Cretaceous Research, 32, 433-446.
Ibrahim, N., Unwin, D., Martill, David, Baidder, L. and Zouhri, S. (2010) A new pterosaur (Pterodactyloidea: Azhdarchidae) from the Upper Cretaceous of Morocco. PloS ONE, 5 (5). e10875
Ibrahim, N. (2007); Berlin shows how natural history can pull the crowds, Nature, 448 (7156): 864.
Published Scientific Abstracts (* indicates speaker) - selection
* McPhee, J., Ibrahim, N., Kao, A., Unwin, D.M., Smith, R., Martill, D.M. (2019) Edentulous pterosaurs from the Cretaceous Kem Kem beds of Southern Morocco: a new genus and species and an upper jaw for Alanqa saharica. Symposium of Vertebrate Palaeontology and Comparative Anatomy Annual Meeting, Isle of Wight, Program and Abstracts.
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* Martill, D.M., Williams, C., Kao, A., Keeble, W., Ibrahim, N. (2019) Internal radial spoke-like trabeculae resist torsion and bending in hyper-elongate azhdarchid pterosaur vertebrae. Symposium of Vertebrate Palaeontology and Comparative Anatomy Annual Meeting, Isle of Wight, Program and Abstracts.
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* Smyth, R., Ibrahim, N., Martill, D.M.M. (2019) How many spinosaur taxa are there in the Kem Kem beds of Morocco. Symposium of Vertebrate Palaeontology and Comparative Anatomy Annual Meeting, Isle of Wight, Program and Abstracts.
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* Bindellini, G., Matarelli, D., Auditore, M., Maganuco, S., Ibrahim, N., Fabbri, M., Martill, D.M., Zouhri, S., Dal Sasso, C. (2019) The crucial role of photogrammetry in reconstructing the foot skeleton of Spinosaurus. Paleodays 2019, XIX Edizione delle Giornata di Paleontologia, Program and Abstracts.
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* Williams, C., Martill, D.M., Kau, A., Keeble, W., Ibrahim, N. (2019) Radial spoke-like trabeculae provide strength in hyper-elongate pterosaur vertebrae. 17th Conference of the European Association of Vertebrate Palaeontologists, Program and Abstracts.
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* Smyth, R.S.H., Martill, D.M., Ibrahim, N. (2019) CT analysis reveals new information on the diversity of small theropods oft he Kem Kem beds, southeastern Morocco. 17th Conference of the European Association of Vertebrate Palaeontologists, Program and Abstracts.
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* Smith, R.E., Martill, D.M., Ibrahim, N. (2019) The taphonomy of pterosaurs from the Cretaceous Kem Kem beds of S.E. Morocco. 17th Conference of the European Association of Vertebrate Palaeontologists, Program and Abstracts.
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* Dahdul, W., Manda, P., Cui, H., Balhoff, J., Dececchi, A., Ibrahim, N., Lapp, H., Mabee, P., Vision, T. (2017) Gold Standard evaluation of machine and human generated annotations of biodiverse phenotypes. Biocuration 2017, Abstracts.
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*Fabbri, M., Benson, R., Pol, D., Orkney, A., Dal Sasso, C., Maganuco, S., Zouhri, S., Ibrahim, N. (2018) Fish-eating habits in spinosaurs are shaped by heterochrony and bone microstructure. 5th International Paleontological Congress, Abstract Book, 2018: 704.
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* Dahdul, W., Manda, P., Cui, H., Balhoff, J., Dececchi, A., Ibrahim, N., Lapp, H., Mabee, P., Vision, T. (2017) Gold Standard evaluation of machine and human generated annotations of biodiverse phenotypes. Biocuration 2017, Abstracts.
*Fabbri, M., Dal Sasso, C., Maganuco, S., Zouhri, S., Martill, D.M., Ibrahim, N. (2017) Bone microstructure and heterochrony shape fish-eating habits in spinosaurs. Journal of Vertebrate Paleontology, Program and Abstracts, 2017: 109.
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*Ibrahim, N., Maganuco, S., Sereno, P.C., Dal Sasso, C., Keillor, T., Martill, D.M., Zouhri, S., Fabbri, M., Auditore, M. 2014. Associated remains of Spinosaurus aegyptiacus, an enormous predatory dinosaur with subaquatic adaptations. Journal of Vertebrate Paleontology, Program and Abstracts, 2014: 151.
Dahdul, W.M., Dececchi, T.A., Ibrahim, N., Lapp, H., Mabee, P. 2014. Moving the mountain: How to transform comparative anatomy into computable anatomy? Evolution 2014, Raleigh, Abstract Volume.
*Conroy, L., Keillor, T., Gruszauskas, N., Ibrahim, N., Sereno, P.C. 2014. From Fossils to Digital Models: CT Scanning and Modeling for Rapid Prototyping and Animation. Digital Specimen 2014, Museum für Naturkunde Berlin, Abstracts of Presentations.
*Ibrahim, N., 2014. A River of Giants. Second International Congress on North African Vertebrate Palaeontology, Ouarzazate, Abstract Volume.
*Ibrahim, N., 2014. Ernst Stromer von Reichenbach: a Founding Father of North African Vertebrate Paleontology. Second International Congress on North African Vertebrate Palaeontology, Ouarzazate, Abstract Volume.
*Mungall, C.J., Balhoff, J., Bastian, F., Blackburn, D., Comte, A., Dahdul, WM., Dececchi, T.A., Ibrahim, N., Lewis, S.E., Mabee, P., Niknejad, A., Haendel, M. 2013. A merger of multi-species anatomy ontologies. Biocuration conference, Cambridge, UK, Abstract Volume.
Mabee, P.M., Dececchi, T.A., Ibrahim, N., Sereno, P.C., Balhoff, J.P. 2013. Aggregation of morphological characters across studies using an ontology-based phenotype approach. Journal of Vertebrate Paleontology, Program and Abstracts, 2013: 51.
Sereno, P.C., *Ibrahim, N., Mabee, P., Vision, T., Lapp, H. 2012. Phenoscape: An anatomical ontology of vertebrates. Journal of Vertebrate Paleontology, Program and Abstracts, 2009: 170-171.
*Zouhri, S., Ibrahim, N., Baidder, L., Martill, D.M., Kaoukaya, A. 2012. ''Kem Kem Beds'': un ''point chaud'' de la paléobiodiversité terrestre au Crétacé superieur basal. Rencontre sur la valorisation et la protection du patrimoine paléontologique (RV3P4), Morocco, book of abstracts.
*Ibrahim, N., Sereno, P.C. 2011. New data on spinosaurids (Dinosauria: Theropoda) from North Africa. Journal of Vertebrate Paleontology, Program and Abstracts, 2011: 130.
*Ibrahim, N. 2010. A unique ancient ecosystem: the theropod dominated Late Cretaceous Kem Kem dinosaur assemblage of South East Morocco. Journal of Vertebrate Paleontology, Program and Abstracts, 2010: 109.
*Ibrahim, N., Unwin, D.M., Zouhri, S., Martill, D.M. 2009. Diversity and evolutionary significance of pterosaurs from the early Late Cretaceous of Morocco. Journal of Vertebrate Paleontology, Program and Abstracts, 2009: 119.
*Martill, D.M., Ibrahim, N., Unwin, D.M., 2009. Pachyostosis in azhdarchoid pterosaur beaks from the Kem Kem Formation (early Late Cretaceous) of SE Morocco. Mesozoic Terrestrial Ecosystems meeting, Teruel, Spain, Abstract volume, p. 75.
*Ibrahim, N., Zouhri, S, Baidder, L. 2009. Fossil vertebrates from the early Late Cretaceous of Morocco. First International Congress on North African Vertebrate Palaeontology, Abstract Volume.
*Ibrahim, N. 2008. Too many theropods? The diversity of predatory dinosaurs in the mid-Cretaceous of Morocco. Symposium of Vertebrate Palaeontology and Comparative Anatomy Annual Meeting, Dublin, Abstract Volume.
*Ibrahim, N., Baidder, L., Dyke, G.J., Jalil, N.-E., Ouanaimi, F., Schouten, R., Zouhri, S. 2007. Vertebrate Palaeontology of the Kem Kem Beds, Morocco, and Palaeoecology of North Africa. Symposium of Vertebrate Palaeontology and Comparative Anatomy Annual Meeting, Glasgow, Abstract Volume.
Schouten, R., Dyke, G.J., Ibrahim, N., Jalil, N.-E., Zouhri, S. 2007. Vertebrate paleontology and geological setting of the Kem Kem Beds, Morocco. Journal of Vertebrate Paleontology, Program and Abstracts, 2007: 142.
*Ibrahim, N. 2007. Review of North African dinosaur faunas during the Cretaceous. First Mediterranean Herpetological Congress, Marrakech, Morocco, Abstract Volume.
*Ibrahim, N. 2007. North African Vertebrate Faunas. Irish Geological Research Meeting, Ulster, Abstract volume.
Popular Books, Articles
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Ibrahim, N., Joger, U., Kosma, R., Beckmann, A. (2018). Spinosaurus – Der rätselhafte Riese. Cargo-Verlag, Schwülper, 76 pp. [Exhibit Guide book, in German]
Ibrahim, N. 2007; Spinosaurs, Prehistoric Times, 83.
Smith, A., Ibrahim, N. 2007; Sticking their necks out – long necks in prehistoric animals,
Rockwatch, 46, 14-15.
Ibrahim, N. 2004; British Dinosaurs, Rockwatch